Premolars compressed, pointed; and the molars with quadrate tuberculated crowns.
The maxillaries are connected with the distal anterior corner of the quadrate by the thin, splint-like jugal and quadratojugal.
The quadrate is invariably a conspicuous bone and movably articulating with the cranium and by a special process with the pterygoid.
The quadrate muscle adjusts the motion, and prevents pressure upon the optic nerve; during the state of relaxation of both muscles the nictitans withdraws through its own elasticity.
A diagnosis covering all the Ratitae (struthio, rhea, casuarius, dromaeus, apteryx and the allied fossils dinornis and aepyornis) would be as follows - (i) terrestrial birds without keel to the sternum, absolutely flightless; (ii) quadrate bone with a single proximal articulating knob; (iii) coracoid and scapula fused together and forming an open angle; (iv) normally without a pygostyle; (v) with an incisura ischiadica; (vi) rhamphotheca compound; (vii) without apteria or bare spaces in the plumage; (viii) with a complete copulatory organ, moved by skeletal muscles.