In the epistle ambo at Salerno and the gospel ambones at Cava and San Giovanni del Toro in Ravello, the columns support segmental arches carrying the ambones; the epistle ambo at Ravello and all those in Rome are raised on solid marble bases.
The regions of this cuticle have a markedly segmental arrangement, and the definite hardened pieces (sclerites) of the exoskeleton are in close contact with one another along linear sutures, or are united by regions of the cuticle which are less chitinous and more membranous, so as to permit freedom of movement.
In the abdominal exoskeleton the segmental structure is very clearly marked, a series of sclerites - dorsal terga and abdominal sterna - being connected by pale, feebly chitinized cuticle, so that considerable freedom of movement between the segments is possible.
In front of the three masses that will form the sub-oesophageal ganglion the mass of cells that is to form the nervous system is very large, and projects on each side; this anterior or " brain " mass consists of three lobes (the prot-, deut-, and tritencephalon of Viallanes and others), each of which might be thought to represent a segmental ganglion.
This has not been demonstrated by an actual following out of the development, but the position of these pieces and the fact that they are (in Limulus) supplied by an independent segmental nerve, favours the view that they may comprise the sternal area of the vanished praegenital somite.