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At the end of the airways are clusters of tiny air sacs.
Special sacs developed from the intersegmental septa lodge the developing ova and sperm.
The walls of the renal sacs are deeply plaited and thrown into ridges.
Spherical sacs, bearing forked spines, described by Williamson under the name of Zygosporites, are frequent, usually in an isolated state.
The sporangia were large pyriform sacs, shortly stalked, and borne in tufts on the branches of the fertile rachis, which developed no lamina.
