Appendages of 1st pair consisting of three segments, completely chelate, without poison gland; of 2nd pair slender, leg-like, tipped with three claws, the basal segment without sterno-coxal process taking no share in mastication, and widely separated from its fellow of the opposite side; 3rd, 4th, 5th and 6th appendages similar in form to the 2nd and to each other.
Owen's description showed this view to be groundless, and he attributed the extraordinary development of the toucan's beak to the need of compensating, by the additional power of mastication thus given, for the absence of any of the grinding structures that are so characteristic of the intestinal tract of vegetable-eating birds - its digestive organs possessing a general simplicity of formation.
When the summits of these are worn by mastication their surfaces present circles of dentine surrounded by a border of enamel, and as attrition proceeds different patterns are produced by the union of the bases of the cusps, a trefoil form being characteristic of some species.
Mastication, and by filtering through wire-gauze filters, is kept warm by a steam-jacket.
Before commencing the mastication it is generally necessary to warm the apparatus by means of steam; but as the operation proceeds the heat produced requires to be moderated by streams of cold water flowing through channels provided for the purpose.