The antlers of females are simple and generally smaller.
It has been noticed at Woburn Abbey that the antlers are shed and replaced twice a year.
This skin, with the skull and antlers, was sent to Paris, where it was described in 1866 by Professor Milne-Edwards.
The antlers have the trez-tine near the small brow-tine, and the palmation beginning near the former.
In the deer of the sambar group, where the antlers never advance beyond a three-tined type, the shedding is frequently, if not invariably, very irregular; but in the majority at least of the species with complex antlers the replacement is annual, the new appendages attaining their full development immediately before the pairing-season.