Such obvious features as the number of segments in the foot and the shape of the feeler were used by the early entomologists for distinguishing the great groups of beetles.
Fielde show that an ant follows her own old track by a scent exercised by the tenth segment of the feeler, recognizes other inmates of her nest by a sense of smell resident in the eleventh segment, is guided to the eggs, maggots and pupae, which she has to tend, by sensation through the eighth and ninth segments, and appreciates the general smell of the nest itself by means of organs in the twelfth segment.
- a, Saw-toothed Grain-Beetle (Silvanus surinamensis); b, pupa; c, larva, magnified 12 times; d, feeler of larva.
I and 25) have the terminal antennal segments pectinate, and so arranged that the comb-like part of the feeler cannot be curled up, while the elytra completely cover the abdomen.
The Passalidae are a tropical family of beetles generally considered to be intermediate between stag-beetles and chafers, the enlarged segments of the feeler being capable of close approximation.