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Physically and physiologically dry habitats, with the accompanying plant communities of sand dunes and sandy heaths with little humus in the soil.
The former is poor in Cruciferae, Caryophyllaceae, Umbelliferae, Primulaceae and Labiatae; but for the occurrence of Calluna in Newfoundland it would have no heaths.
They are linked together by the presence of Proteaceae and of Epacrideae, which take the place of the nearly allied heaths in South Africa.
The absence of the oak and of all heaths east of the Ural may be noticed.
Heaths, generally somewhat rare in South Africa outside the Cape peninsula, are abundant in Basutoland.